The Transformation of Energy

The transformations of energy in an ecosystem begin first with the input of energy from the sun. Energy from the sun is captured by the process of photosynthesis. Carbon dioxide is combined with hydrogen (derived from the splitting of water molecules) to produce carbohydrates (CHO). Energy is stored in the high energy bonds of adenosine triphosphate.

The prophet Isaah said "all flesh is grass", earning him the title of first ecologist, because virtually all energy available to organisms originates in plants. Because it is the first step in the production of energy for living things, it is called primary production. Herbivores obtain their energy by consuming plants or plant products,carnivores eat herbivores, and detritivores consume the droppings and carcasses of us all. 

Figure 01

Figure 01 portrays a simple food chain, in which energy from the sun, captured by plant photosynthesis, flows fromtrophic level to trophic level via the food chain. A trophic level is composed of organisms that make a living in the same way, that is they are all primary producers (plants), primary consumers (herbivores) or secondary consumers (carnivores). Dead tissue and waste products are produced at all levels. Scavengers, detritivores, and decomposers collectively account for the use of all such "waste" -- consumers of carcasses and fallen leaves may be other animals, such as crows and beetles, but ultimately it is the microbes that finish the job of decomposition. Not surprisingly, the amount of primary production varies a great deal from place to place, due to differences in the amount of solar radiation and the availability of nutrients and water.

Usually when we think of food chains we visualize green plants, herbivores, and so on. These are referred to as grazer food chains, because living plants are directly consumed. In many circumstances the principal energy input is not green plants but dead organic matter. These are called detritus food chains. Examples include the forest floor or a woodland stream in a forested area, a salt marsh, and most obviously, the ocean floor in very deep areas where all sunlight is extinguished 1000's of meters above. 

Finally, although we have been talking about food chains, in reality the organization of biological systems is much more complicated than can be represented by a simple "chain". There are many food links and chains in an ecosystem, and we refer to all of these linkages as a food web. Food webs can be very complicated, where it appears that "everything is connected to everything else", and it is important to understand what are the most important linkages in any particular food web. 

Biogeochemistry:

How can we study which of these linkages in a food web are most important? One obvious way is to study the flow of energy or the cycling of elements. For example, the cycling of elements is controlled in part by organisms, which store or transform elements, and in part by the chemistry and geology of the natural world. The term Biogeochemistry is defined as the study of how living systems influence, and are controlled by, the geology and chemistry of the earth. Thus biogeochemistry encompasses many aspects of the abiotic and biotic world that we live in.

There are several main principles and tools that biogeochemists use to study earth systems. Most of the major environmental problems that we face in our world toady can be analyzed using biogeochemical principles and tools. These problems include global warming, acid rain, environmental pollution, and increasing greenhouse gases. The principles and tools that we use can be broken down into 3 major components: element ratios, mass balance, and element cycling.

Source: http://www.globalchange.umich.edu/globalchange1/current/lectures/kling/ecosystem/ecosystem.html